The availability or lack of nutrients shapes ecosystems in fundamental ways. From forest productivity to soil fertility, from the diversity of animals to the composition of microbial communities, nutrient cycling and limitation are the basic mechanisms underlying ecosystem ecology. In this book, Peter Vitousek builds on over twenty years of research in Hawai'i to evaluate the controls and consequences of variation in nutrient availability and limitation. Integrating research from geochemistry, pedology, atmospheric chemistry, ecophysiology, and ecology, Vitousek addresses fundamental questions: How do the cycles of different elements interact? How do biological processes operating in minutes or hours interact with geochemical processes operating over millions of years? How does biological diversity interact with nutrient cycling and limitation in ecosystems? The Hawaiian Islands provide the author with an excellent model system for answering these questions as he integrates across levels of biological organization. He evaluates the connections between plant nutrient use efficiency, nutrient cycling and limitation within ecosystems, and nutrient input-output budgets of ecosystems. This book makes use of the Hawaiian ecosystems to explore the mechanisms that shape productivity and diversity in ecosystems throughout the world. It will be essential reading for all ecologists and environmental scientists.

In all fields of science today, data are collected and theories are developed and published faster than scientists can keep up with, let alone thoroughly digest. In ecology the fact that practitioners tend to be divided between such subdisciplines as aquatic and terrestrial ecology, as well as between popula tion, community, and ecosystem ecology, makes it even harder for them to keep up with all relevant research. Ecologists specializing in one sub discipline are not always aware of progress in another subdiscipline that relates to their own. Syntheses are frequently needed that pull together large bodies of information and organize them in ways that makes them more coherent, and thus more understandable. I have tried to perform this task of integration for the subject area that encompasses the interrelationships between the dynamics of ecological food webs and the cycling of nutrients. I believe this area cuts across many of the subdisciplines of ecology and is pivotal to our progress in understanding ecosystems and in dealing with human impacts on the environment. Many current ecological problems involve human disturbances of both food webs and the nutrients that cycle through them. Little progress can be made towards elucidating the complex feedback relations inherent in the study of nutrient cycles in ecological systems without the tools of mathematics and computer modelling. These tools are therefore liberally used throughout the book.

The long-term productivity of forest ecosystems depends on the cycling of nutrients. The effect of carbon dioxide fertilization on forest productivity may ultimately be limited by the rate of nutrient cycling. Contemporary and future disturbances such as climatic warming, N-deposition, deforestation, short rotation sylviculture, fire (both wild and controlled), and the invasion of exotic species all place strains on the integrity of ecosystem nutrient cycling. Global differences in climate, soils, and species make it difficult to extrapolate even a single important study worldwide. Despite advances in the understanding of nutrient cycling and carbon production in forests, many questions remain. The chapters in this volume reflect many contemporary research priorities. The thirteen studies in this volume are arranged in the following subject groups: • N and P resorption from foliage worldwide, along chronosequences and along elevation gradients; • Litter production and decomposition; • N and P stoichiometry as affected by N deposition, geographic gradients, species changes, and ecosystem restoration; • Effects of N and P addition on understory biomass, litter, and soil; • Effects of burning on soil nutrients; • Effects of N addition on soil fauna.

It is a well-known fact that eutrophication of coastal waters causes significant changes in the species composition of the primary producers. Usually a shift from an ecosystem dominated by sea grasses or large brown algae to an ecosystem dominated by fast-growing green algae or phytoplankton is observed. While this shift has been documented in a number of research papers and books, the consequences of this shift are less well known. This book focuses on the consequences of such changes for nutrient cycling. The aim is to investigate how different types of primary producers influence nutrient cycling in coastal marine waters, and how nutrient cycling changes qualitatively and quantitatively as a consequence of the changes in the primary producer community caused by eutrophication. The various chapters address specific ecological processes such as grazing, decomposition, burial and export of biomass from the ecosystem. The book is intended for researchers and professionals working in the field of coastal marine ecology and estuarine ecology and for advanced students in this field.

African papyrus (Cyperus papyrus L.) wetlands provide water, food and materials to millions of people, and perform important landscape functions such as water and nutrient storage, habitat provision for fish, birds and other wildlife. They are also an integral part of the culture of African wetland communities. With an increasing demand for food, papyrus wetlands are at risk of conversion to agriculture and losing these ecosystem services. Combining increased agricultural production with wetland conservation is urgently needed. The research presented in this book consisted of two parts. First, field experiments investigated nitrogen and phosphorus retention, showing that papyrus grows faster with disturbance from human activities or flooding, but produces less biomass and stores less nutrients. Then, a dynamic simulation model (Papyrus Simulator) based on the hydrological and ecological wetland processes showed that assimilation, mortality, decay, re-translocation, nutrient inflow and soil porosity were the most influential factors. The model demonstrated that controlled harvesting can increase nutrient retention by up to 40%, but overharvesting leads to the release of nutrients. These findings can help determining optimum harvesting strategies for constructed and natural wetlands, and contribute to the quantification of ecosystem services and an evidence-based adaptive management approach for African wetland landscapes.

The PELAG 1996 Symposium Proceedings provides the reader with the latest advances in the study of planktonic cycling of matter and energy, placing a strong emphasis on the effects of eutrophication on these processes. This book covers a wide range of topics in the field, including: Nutrient limitation of phytoplankton growth Nutrient cycles in the planktonic food web DOM sources, composition, and uptake Resource limitation vs. shaping of the food web by grazing Spatio-temporal variability: coupling of physical and biological processes Processes controlling sinking losses from the pelagic system Planktonic food web modelling . The book should be of interest to everybody involved in planktonic ecosystem research, from the advanced student to the distinguished scientist. This volume brings to the reader the expertise of internationally renowned authors on the main issues of today's ecological plankton research.

In the past 10 years, there has been a resurgence in interest in soil management and conservation of the soil resource. With the knowledge we have accumulated in the past 100 years, there is the possibility of developing new and innovative ways of effectively managing the soil. The emphasis on sustainable agriculture requires that we understand how to utilize the soil as a viable living resource. To meet the world demand for food within the next 50 years requires a healthy and strong soil resource which can sustain production. With the dedication of the National Soil Tilth Laboratory in 1989, it was decided that one contribution the laboratory could make to agri cultural science would be to foster an exchange of information on soil management. The focus of that interchange centers on long-term soil management. If we are to fulfill the goals of sustainable agriculture, environmental quality, and feeding the world, there will have to be an increased understanding of how to effectively manage the soil. Long-term soil management requires integrated and interdisciplinary research to bring all of the information together in terms which would be applicable to all soils. To accomplish this goal a workshop is held each year, with the exchange of information focusing on a single topic within the framework of developing effective strategies for long-term soil management. With the forum to focus on an individual theme each year, the theme for the initial workshop was "Limitations to Plant Root Growth.

This book presents the latest information on tropical tree physiology, making it a valuable research tool for a wide variety of researchers. It is also of general interest to ecologists (e.g. Ecological Society of America; > 3000 or 4000 members at annual meeting), physiologists (e.g. American Society of Plant Biologists; > 2,000 members at annual meeting), and tropical biologists (e.g. Association for Tropical Biology and Conservation, ATBC; > 500 members at annual meeting). (American Geophysical Union(AGU), > 20000 members at annual meeting). Since plant physiology is taught at every university that offers a life sciences, forestry or agricultural program, and physiology is a focus at research institutes and agencies worldwide, the book is a must-have for university and research institution libraries.

Inputs of nitrogen to terrestrial and aquatic ecosystems have increased several-fold over the last one hundred and fifty years, with the steepest increases during the last four decades. The expansion of fertilizer manu facture and use, the increase in fossil fuel combustion, the intensification of animal husbandry, and widespread cultivation of N2 fixing crops have all contributed to the dramatic increase in N inputs. The increase has been most rapid in Northern Hemisphere (NH) temperate ecosystems, but presently subtropical and tropical regions of Asia are also experiencing an explosive increase in N inputs to terrestrial ecosystems (W. Chameides, pers. comm. ; Galloway et al. 1996). Projected increases in N deposition for these trop ical and subtropical regions, with a high natural background of N inputs, exceed increases projected for temperate and arctic regions (Cleveland et al. submitted; Galloway et al. 1994; Holland & Lamarque 1997a). Compared to biological N fixation, N deposition is becoming a proportionately greater source of N to terrestrial and aquatic ecosystems worldwide (Vitousek et al. 1997). 6 The nitrogen contained in the atmosphere as N , 3. 9 * 10 Tg (Tg = 2 12 10 g), is the largest reservoir of N in the Earth system (Warneck 1988). However, this paper focuses on the nitrogen emissions and deposition that have been transformed from N2 into reactive forms that are biologically avail able (e. g. Vitousek et al. 1997).