When we consider the main object of forestry, the tree, it immediately becomes clear why experimental population geneticists have been so hesitant in making this object a primary concern of their research. Trees are very long-living organisms with generation intervals frequently exceeding those of their investigators by multiples. They virtually exclude, therefore, application of the classical methods of population genetics since these are based on observing genetic structures over generations. This situation, where the limits set to observation are so severe, particularly requires close cooperation between theory and experiment. It also requires careful consideration of results obtained for organisms other than trees, in order to gain additional insights by comparing the results for trees with those for other organisms. Yet, the greatest challenge to population and ecological genetics probably originates from the fact that forests are very likely to be the most complex ecosystems of all, even in some cases where they are subject to intense management. This complexity, which equally comprises biotic and abiotic factors varying both in time and space, makes extremely high demands on the adaptational capacity and thus flexibility of the carriers of such an ecosystem. Longevity combined with immobility during the vegetative phase, however, appears to contradict the obvious necessity of adaptational flexibility in forest tree populations when compared with short lived and/or mobile organisms.

Tropical climates, which occur between 23°30'N and S latitude (Jacob 1988), encompass a wide variety of plant communities (Hartshorn 1983, 1988), many of which are diverse in their woody floras. Within this geographic region, temperature and the amount and seasonality of rainfall define habitat types (UNESCO 1978). The F AO has estimated that there 1 are about 19 million km of potentially forested area in the global tropics, of which 58% were estimated to still be in closed forest in the mid-1970s (Sommers 1976; UNESCO 1978). Of this potentially forested region, 42% is categorized as dry forest lifezone, 33% is tropical moist forest, and 25% is wet or rain forest (Lugo 1988). The species diversity of these tropical habitats is very high. Raven (1976, in Mooney 1988) estimated that 65% of the 250,000 or more plant species of the earth are found in tropical regions. Of this floristic assemblage, a large fraction are woody species. In the well-collected tropical moist forest of Barro Colorado Island, Panama, 39. 7% (481 of 1212 species) of the native phanerogams are woody, arborescent species (Croat 1978). Another 21. 9% are woody vines and lianas. Southeast Asian Dipterocarp forests may contain 120-200 species of trees per hectare (Whitmore 1984), and recent surveys in upper Amazonia re corded from 89 to 283 woody species ~ 10 cm dbh per hectare (Gentry 1988). Tropical communities thus represent a global woody flora of significant scope.

Air pollutants provide environmental conditions that drastically differ in many respects from those to which forest trees are naturally adapted. Leading experts in the field here consider these questions of immediate relevance arising from the changing environment: (1) Do air pollutants introduce effects of selection that differ from those known for populations that are not subject to such stress conditions? (2) If air pollutants introduce quantitatively or even qualitatively novel selective effects, which consequences might arise from the adaptation of forest tree populations to the present conditions as well as for the preservation of adaptability to future conditions? In addition to these questions, concepts for preservation of genetic resources are discussed.

This book, which contains 20 chapters, integrates the varied subdisciplines of genetics and their applications in gene conservation, tree improvement and biotechnology. Topics covered include: genetic variation in natural forests, the application of genetics in tree improvement and breeding programmes, and genomic sequences and molecular technologies. This book will be a valuable resource for students, scientists and professionals in the plant sciences, especially forest geneticists, tree breeders, forest managers and other natural resource specialists.

During the 4th International Consultation on Forest Genetics and Tree Building, held in 1998 in Beijing, China, leading scientists were invited to review past achievements, to redefine the role of forest genetics and breeding in contemporary forestry, and to set priorities for future research and development. On the basis of the invited presentations, the volume summarizes the state of knowledge in various fields, such as the impact of forest management and of changing environmental conditions on genetic resources, and the value of genetic markers as indicators for adaptational potential, as well as the tasks of conservation. Detailed reports from the different continents provide a comprehensive diagnosis of the global situation of forest genetics and tree breeding research. The book offers not only an overview of contemporary trends and expected future developments, but also identifies current main problems in funding and cooperation. It may prove therefore useful not only for scientists, university lecturers and advanced students in the field of forestry, ecology and conservation biology, but also for decision makers and managers in companies and conservation organizations.

This book is a printed edition of the Special Issue "Genetics and Genomics of Forest Trees" that was published in Forests