The Biology of Sharks and Rays is a comprehensive resource on the biological and physiological characteristics of the cartilaginous fishes: sharks, rays, and chimaeras. In sixteen chapters, organized by theme, A. Peter Klimley covers a broad spectrum of topics, including taxonomy, morphology, ecology, and physiology. For example, he explains the body design of sharks and why the ridged, toothlike denticles that cover their entire bodies are present on only part of the rays’ bodies and are absent from those of chimaeras. Another chapter explores the anatomy of the jaws and the role of the muscles and teeth in jaw extension, seizure, and handling of prey. The chapters are richly illustrated with pictures of sharks, diagrams of sensory organs, drawings of the body postures of sharks during threat and reproductive displays, and maps showing the extent of the species’ foraging range and long-distance migrations. Each chapter commences with an anecdote from the author about his own personal experience with the topic, followed by thought-provoking questions and a list of recommended readings in the scientific literature. The book will be a useful textbook for advanced ichthyology students as well as an encyclopedic source for those seeking a greater understanding of these fascinating creatures.

This volume had its origin in a symposium on the Reproduction and Development of Cartilaginous Fishes that was held at the annual meetings of the American Elasmobranch Society and the American Society of Ichthyologists and Herpetologists in Charleston, South Carolina in 1990. The cartilaginous fishes, class Chondrichthyes, are a large and diverse group of fishes that include approximately 900 to 1100 living species of sharks, skates, rays and ratfishes. Throughout their history, which dates back at least 400 million years, they have been a successful major component of the marine ecosystem. The chondrichthyan fishes occupy a pivotal position in comparative and evolutionary studies of vertebrate reproduction and development. They are the oldest surviving group of jawed vertebrates and they possess both the adult vertebrate Bauplan and the vertebrate program of embryonic development. The major features of the female reproductive system, including its embryonic origin, structure, physiological function, and biochemistry, apparently were established early in vertebrate evolution and are fully developed in chondrichthyan fishes. These features of the female reproductive system have been retained during the evolution of the other classes of vertebrates. Much the same can be said for the male reproductive system. Moreover, viviparity, placental nourishment of developing embryos, and the hormonal regulation of these events made an initial appearance in this group. The 22 articles presented in this volume bring together a wide variety of complementary research by investigators from seven countries, allowing us to broaden the scope and implications of our studies while identifying opportunities for future research. The appearance of a volume on the reproduction and development of cartilaginous fishes is quite opportune. The continued existence of these fishes, which survived the great extinction events of Earth's history, is now threatened by overexploitation unless immediate steps for their conservation are undertaken. Knowledge of their reproduction and development not only is an end in itself, but is of critical importance in devising successful conservation and resource management strategies.

Successor to the classic work in shark studies, The Elasmobranch Fishes by John Franklin Daniel (first published 1922, revised 1928 and 1934), Sharks, Skates, and Rays provides a comprehensive and up-to-date overview of elasmobranch morphology. Coverage has been expanded from anatomy to include modern information on physiology and biochemistry. The new volume also provides equal treatment for skates and rays. The authors present general introductory material for the relative novice but also review the latest technical citations, making the book a valuable primary reference resource. More than 200 illustrations supplement the text.

This volume constitutes a series of invited chapters based on presentations given at an International Conference on the Sensory Biology of Aquatic Animals held June 24-28, 1985 at the Mote Marine Laboratory in Sarasota, Florida. The immediate purpose of the conference was to spark an exchange of ideas, concepts, and techniques among investigators concerned with the different sensory modalities employed by a wide variety of animal species in extracting information from the aquatic environment. By necessity, most investigators of sensory biology are specialists in one sensory system: different stimulus modalities require different methods of stimulus control and, generally, different animal models. Yet, it is clear that all sensory systems have principles in common, such as stimulus filtering by peripheral structures, tuning of receptor cells, signal-to-noise ratios, adaption and disadaptation, and effective dynamic range. Other features, such as hormonal and efferent neural control, circadian reorganization, and receptor recycling are known in some and not in other senses. The conference afforded an increased awareness of new discoveries in other sensory systems that has effectively inspired a fresh look by the various participants at their own area of specialization to see whether or not similar principles apply. This inspiration was found not only in theoretical issues, but equally in techniques and methods of approach. The myopy of sensory specialization was broken in one unexpected way by showing limitations of individual sense organs and their integration within each organism. For instance, studying vision, one generally chooses a visual animal as a model.

Virtually every area of research associated with sharks and their relatives has been strongly impacted by the revolutionary growth in technology. The questions we can now ask are very different than those reported even two decades ago. Modern immunological and genetic techniques, satellite telemetry and archival tagging, modern phylogenetic analysis, GIS, and bomb dating, are just a few of the techniques and procedures that have become a part of our investigative lexicon. A modern synthesis of the biology of Chondrichthyans, Biology of Sharks and Their Relatives, Second Edition discusses significant advances in the development and application of new molecular techniques to the understanding of the phylogenetic relationships among and between these groups. The book considers the effect of global changes on the status of sharks and their relatives, and how advances in technology and analytical techniques have changed not only how we approach problem solving and scientific investigations, but how we formulate questions. The book also introduces applications of new and novel laboratory devices, techniques, and field instruments. This second edition of the award winning and groundbreaking original exploration of the fundamental elements of the taxonomy, systematics, physiology, and ecology of sharks, skates, rays, and chimera, presents cohesive and integrated coverage of key topics and discusses technological advances used in modern shark research. Offering a well-rounded picture for students and researchers, and far above competitors in scope and research, this new volume holds a wealth of data on the current status of Chondrichthyan research and provides the basis and springboard for original research. Cover photo by Justin Gilligan

Biology of Sharks and Their Relatives is an award-winning and groundbreaking exploration of the fundamental elements of the taxonomy, systematics, physiology, and ecology of sharks, skates, rays, and chimera. This edition presents current research as well as traditional models, to provide future researchers with solid historical foundations in shark research as well as presenting current trends from which to develop new frontiers in their own work. Traditional areas of study such as age and growth, reproduction, taxonomy and systematics, sensory biology, and ecology are updated with contemporary research that incorporates emerging techniques including molecular genetics, exploratory techniques in artificial insemination, and the rapidly expanding fields of satellite tracking, remote sensing, accelerometry, and imaging. With two new editors and 90 contributors from the US, UK, South Africa, Portugal, France, Canada, New Zealand, Australia, India, Palau, United Arab Emirates, Micronesia, Sweden, Argentina, Indonesia, Cameroon, and the Netherlands, this third edition is the most global and comprehensive yet. It adds six new chapters representing extensive studies of health, stress, disease and pathology, and social structure, and continues to explore elasmobranch ecological roles and interactions with their habitats. The book concludes with a comprehensive review of conservation policies, management, and strategies, as well as consideration of the potential effects of impending climate change. Presenting cohesive and integrated coverage of key topics and discussing technological advances used in modern shark research, this revised edition offers a well-rounded picture for students and researchers.

The elasmobranch fishes include the living sharks, skates and rays that are important members of nearly all marine ecosystems. Their large size, secretive behavior, and wide-ranging habits make them difficult to observe in the field or to maintain in captivity. Consequently, little is known about their natural behavior and how it is mediated by their sensory systems. This volume is dedicated to the scientific contributions and memory of Donald Nelson, a pioneer in the study of shark behavior, sensory biology, and remote instrumentation. The two opening papers review Don Nelson's unique scientific accomplishments and provide insight into his strong bias towards study of animals in the field. These are followed by 14 scientific papers on elasmobranch behavior, sensory biology, and current monitoring technologies. The papers on elasmobranch sensory biology and behavior address questions on hearing, the lateral line, electroreception, the brain, orientation behavior, chemical irritants, feeding, and reproduction. The latter section of the volume presents papers on conventional tagging techniques, ultrasonic telemetry, physiological telemetry, remote monitoring techniques, archival tagging and satellite tagging. The intent of this volume is to familiarize both new and established scientists with the sensory biology and behavior of sharks and rays, and to encourage further behavioral research on these animals in their natural environment.

Fish comprise more than 50% of all living vertebrates and are found in a wide range of highly diverse habitats like the deep sea, the shoreline, tide pools, tropical streams and sweetwater ponds. During evolution, the senses of fish have adapted to the physical conditions of the environment in which different species live. As a result, the senses of fish exhibit a remarkable diversity that allows different species to deal with the physical constraints imposed by their habitat. In addition, fish have evolved several `new' sensory systems that are unique to the aquatic environment. In this book, examples of adaptation and refinement are given for six sensory systems: The visual system, The auditory system, The olfactory system, The mechanosensory lateral line system, The taste system, The electrosensory system. In each case, the environmental conditions under which a particular group of fish lives are analyzed. This is followed by a description of morphology and physiology of the sensory system and by an evaluation of its perceptional capabilities. Finally, the sensory adaptations to the particular conditions that prevail in the habitat of a species are highlighted. The various examples from different groups of fish presented in this book demonstrate the impressive capability of fish sensory systems to effectively overcome physical problems imposed by the environment.